Quarterly report pursuant to Section 13 or 15(d)

Collaborative and Other Agreements - NIAID - Additional Information (Details)

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Collaborative and Other Agreements - NIAID - Additional Information (Details) - USD ($)
1 Months Ended 3 Months Ended 9 Months Ended
Oct. 31, 2011
Sep. 30, 2016
Sep. 30, 2015
Sep. 30, 2016
Sep. 30, 2015
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]          
Recognition of revenue   $ 205,000 $ 1,429,000 $ 1,844,000 $ 5,412,000
National Institute of Allergy and Infectious Diseases "NIAID" [Member]          
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]          
Total contract amount awarded $ 28,000,000        
Contractual term 5 years        
Recognition of revenue   $ 0 $ 1,200,000 $ 1,100,000 $ 4,100,000