Quarterly report pursuant to Section 13 or 15(d)

Collaborative, Licensing and Other Arrangements - NIAID - Additional Information (Details)

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Collaborative, Licensing and Other Arrangements - NIAID - Additional Information (Details) - USD ($)
1 Months Ended 3 Months Ended
Oct. 31, 2011
Mar. 31, 2017
Mar. 31, 2016
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]      
Recognition of revenue   $ 110,000 $ 1,471,000
National Institute of Allergy and Infectious Diseases "NIAID" [Member] | Arrangement with Governmental Agency 1 [Member]      
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]      
Total contract amount awarded $ 28,000,000    
Contractual term 5 years    
Recognition of revenue   $ 0 $ 1,100,000