Quarterly report pursuant to Section 13 or 15(d)

Stock-based Compensation - Stock-based Compensation Expense (Details)

v3.7.0.1
Stock-based Compensation - Stock-based Compensation Expense (Details) - USD ($)
$ in Thousands
3 Months Ended 6 Months Ended
Jun. 30, 2017
Jun. 30, 2016
Jun. 30, 2017
Jun. 30, 2016
Employee Service Share-based Compensation, Allocation of Recognized Period Costs [Line Items]        
Total stock-based compensation expense $ 1,854 $ 2,165 $ 2,855 $ 4,471
Research and Development [Member]        
Employee Service Share-based Compensation, Allocation of Recognized Period Costs [Line Items]        
Total stock-based compensation expense 231 857 672 1,994
General and Administrative [Member]        
Employee Service Share-based Compensation, Allocation of Recognized Period Costs [Line Items]        
Total stock-based compensation expense $ 1,623 $ 1,308 $ 2,183 $ 2,477